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As Gambit62 says, B12 is also reserved in the liver

Sources Boston Concierge Orthopedics

"Sensible Drinking Guidelines" (PDF)

This is reflected in the UK guidelines, the absence of a raised MCV cannot be used to exclude the need for cobalamin testing because neurological impairment occurs with a normal MCV in 25% of cases

Growth Factors and Growth Factor Modulators, including, but not limited to: Fibroblast growth factors (FGFs) Hepatocyte growth factor (HGF) Insulin-like growth factor-1 (IGF-1) and its analogues Mechano growth factors (MGFs) Platelet-derived growth factor (PDGF) Thymosin- 4 and its derivatives e.g